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2013 Mar;154(5):681-5

Paracellular intestinal permeability can be assessed in vivo by using different molecules (e.g., sugars, polyethylene glycols, 51 Cr-EDTA) and ex vivo in Ussing chambers combining electrophysiology and probes of different molecular sizes
See these diagrams: Source: Clinical Procedures for Safer Patient Care by Doyle & McCutcheon
